Původní popis
Gilbert, C. H. & E. C. Starks. 1904. The fishes of Panama Bay. Mem. Calif. Acad. Sci. v. 4: 1-304, Pls. 1-33.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.