Původní popis
Heller, E. & Snodgrass, R. E. 1903. Papers from the Hopkins Stanford Galapagos expedition, 1898-1899. XV. New fishes. Proceedings of the Washington Academy of Science. v. 5: 189-229, Pls. 2-20.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.