Původní popis
Fowler, H. W. 1930. The freshwater fishes obtained by the Gray African Expedition--1929. With notes on other species in the Academy collection. Proc. Acad. Nat. Sci. Phila. v. 82: 27-83.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.