Původní popis
Gilbert, C. R. & Kelso, D. P. 1971. Fishes of the Tortuguero area, Caribbean Costa Rica. Bulletin of the Florida State Museum, Biological Sciences v. 16 (no. 1): 1-54.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.