Původní popis
Clark, H. W. 1936. New and noteworthy fishes. The Templeton Crocker expedition of the California Academy of Sciences, 1932. Proceedings of the California Academy of Sciences (Series 4) v. 21 (no. 29): 383-396.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.