Původní popis
Jordan, D. S. & J. O. Snyder. 1901. List of fishes collected in 1883 and 1885 by Pierre Louis Jouy and preserved in the United States National Museum, with descriptions of six new species. Proc. U. S. Natl. Mus. v. 23 (no. 1235): 739-769, Pls. 31-38.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.