Čeleď: Salmonidae Jarocki / Schinz, 1822 — lososovití
Rod: Thymallus Linck, 1790
Knizhin, I.B., Antonov, A.L., Safronov, S.N. & Weiss, S.J. 2007. New Species of Grayling Thymallus tugarinae sp. nova (Thymallidae) from the Amur River Basin. Journal of Ichthyology, 47 (2): 123-139.
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.