Původní popis
Eigenmann, C. H., W. L. McAtee & D. P. Ward 1907. On further collections of fishes from Paraguay. Ann. Carnegie Mus. v. 4 (no. 2): 110-157, Pls. 31-45.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.