Původní popis
Jordan, D. S. & H. W. Fowler. 1903. A review of the cyprinoid fishes of Japan. Proceedings of the United States National Museum v. 26 (no. 1334): 811-862.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.