Původní popis
Fowler, H. W. 1935. South African fishes received from Mr. H. W. Bell-Marley in 1935. Proceedings of the Academy of Natural Sciences of Philadelphia v. 87: 361-408.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.