Původní popis
Regan, C. T. 1913. The fishes of the San Juan River, Colombia. Ann. Mag. Nat. Hist. (Ser. 8) v. 12 (no. 71): 462-473.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.