Původní popis
Meek, S. E. & Hildebrand, S. F. 1916. The fishes of the fresh waters of Panama. Field Museum of Natural History, Publications, Zoölogical Series v. 10 (no. 15): 1-374, Pls. 6-32.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.