Původní popis
Kamohara, T. 1936. Supplementary note on the fishes collected in the vicinity of Kôchi-shi (IX). Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 48 (no. 6): 306-311.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.