Původní popis
Fowler, H. W. 1934. Fishes obtained by Mr. H. W. Bell-Marley chiefly in Natal and Zululand in 1929 to 1932. Proceedings of the Academy of Natural Sciences of Philadelphia v. 86: 405-514.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.