Původní popis
Smith, H. M. & Pope, T. E. B. 1906. List of fishes collected in Japan in 1903, with descriptions of new genera and species. Proceedings of the United States National Museum v. 31 (no. 1489): 459-499.
Použitá literatura
Fricke, R., Eschmeyer, W. N. & Van der Laan, R. (eds). 2024. Eschmeyer's catalog of fishes: Genera, species, references. World Wide Web.