Původní popis
Jordan, D. S. & Snyder, J. O. 1901. A review of the gobioid fishes of Japan, with descriptions of twenty-one new species. Proceedings of the United States National Museum v. 24 (no. 1244): 33-132.
Jordan, D. S. & Snyder, J. O. 1901. A review of the gobioid fishes of Japan, with descriptions of twenty-one new species. Proceedings of the United States National Museum v. 24 (no. 1244): 33-132.
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