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Jordan D. S. & Starks E. C. 1901. A review of the atherine fishes of Japan. Proc. U. S. Natl. Mus. v. 24 (no. 1250). 199-206.
Clark H. W. 1937. New fishes from the Templeton Crocker expedition of 1934-35. Copeia 1937 (no. 2). 88-91.